↑ 1.01.11.2Eigenmann, R.S. (1912) The freshwater fishes of British Guiana, including a study of the ecological grouping of species, and the relation of the fauna of the plateau to that of the lowlands., Mem. Carneg. Mus. 5:578 p.
↑ 2.02.12.2Kullander, S.O. (1986) Cichlid fishes of the Amazon River drainage of Peru., Department of Vertebrate Zoology, Research Division, Swedish Museum of Natural History, Stockholm, Sweden, 394 p.