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↑Endrödi S. (1955) Die Erscheinungsformen der hungarischen Lamellicornia-Arten, Folia Entomologica Hungarica, Budapest 8:45-54
↑ 3.03.1Matsumura S. (1937) The Onthophagid-insects from Korea with descriptions of new species, Insecta matsumurana. Entomological Museum Hokkaido University. Sapporo 12:1-6
↑Reitter E. (1893) Bestimmungs-Tabelle der Lucaniden und coprophagen Lamellicornen des palaearctischen Faunengebietes, Verhandlungen des Naturforschenden Vereins, Brunn (1892) 30:3-230