↑Matsumura S. (1937) New Onthophagus-species in Japan with a tabular key, Insecta matsumurana. Entomological Museum Hokkaido University. Sapporo 11:150-169
↑Gillet J.J.E. (1924) Descriptions d'Onthophagus nouveaux d'Asie tropicale, Annales de la Société scientifique de Bruxelles 44:66-70
↑ 3.03.1Bisby F.A., Roskov Y.R., Orrell T.M., Nicolson D., Paglinawan L.E., Bailly N., Kirk P.M., Bourgoin T., Baillargeon G., Ouvrard D. (ed.) (2011). "Species 2000 & ITIS Catalogue of Life: 2011 Annual Checklist". Species 2000: Reading, UK. Ginkuhà 24 Septyembre 2012.CS1 maint: multiple names: authors list (link) CS1 maint: extra text: authors list (link)
↑Scarabs: World Scarabaeidae Database. Schoolmeesters P., 30 Mayo 2011